| Reference no. | Jefferson, T.A., S. Leatherwood and M.A. Webber, 1993 |
| Ecosystem | Sunda Shelf/Java Sea |
| Status | native |
| Abundance | |
| Lifestage | adults |
| Regional database | |
| Remarks | Known from Borneo (Ref. 87060), particularly in Sarawak (Refs. 85208, 104595), on a beach near the Lutong river (Ref. 87068). C: Ref. 84562; 85208, 104595; M: Ref. 1394; O: Ref. 87068; R: Ref. 87060. |