| Reference no. | Mutlu, E., F. Bingel, A.C. Gücü, V.V. Melnikov, U. Niermann, N.A. Ostra and V.E. Zaika, 1994 |
| Ecosystem | Black Sea - province |
| Status | native |
| Abundance | |
| Lifestage | adults |
| Regional database | |
| Remarks | Found in water column above anoxic zone below maximal pycnocline at depths between 60 to 170 m during the day and migrates to the surface at night (Ref. 3031). Also in Refs. 1656, 96408. |