| Reference no. | Enticott, J. and D. Tipling, 1997 |
| Ecosystem | Temperate Northern Pacific |
| Status | native |
| Abundance | |
| Lifestage | adults |
| Regional database | |
| Remarks | Breeds colonially on steep inaccessible slopes and offshore islands from Nemuro Peninsula, Hokkaido (Ref. 84934). C: Ref. 84934. |