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(English) 1) Water circulation in lakes in spring and autumn leading to an equalisation of the temperature regime throughout the water column and transport of nutrients into the water column, 2) extinction of a species within a given area and its replacement by another, 3) part of a population that is lost by death and emigration and replaced by recruitment per unit time, 4) the ratio of productive energy flow to standing crop biomass in a community or ecosystem.

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