| Main Ref. | Komai, T., 1999 |
| Mode | protandry |
| Fertilization | |
| Spawning Frequency | |
| Batch Spawner | No |
| Reproductive Guild | |
| Description of life cycle and mating behavior | Metamorphosis finishes by late June, growth was rapid until October where males became sexually mature. They remain as males by the second autumn, sex change occurs in the third spring. Spawning starts in October, egg hatches by March and April (Ref. 82562). |
| Search for more references on reproduction | Scirus |