| Kutipan | Conand, C., 1998 Holothurians (sea cucumbers, Class Holothuroidea). p. 1157-1190. In Carpenter, K.E. and V.H. Niem (eds.) FAO Species Identification Guide for Fishery Purposes. The Living Marine Resources of the Western Central Pacific. Vol. 2. Cephalopods, crustaceans, holothurians and sharks. FAO Rome. |
| Acuan | 122 Conand, C., 1998 |
| Halaman | 1176 |
| Bernama Digunakan sebagai Valid | Holothuria atra |
| Lokalitas | |
| Penyebaran |